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Jüri Engelbrecht, Kert Tamm, and Tanel Peets
Centre for Nonlinear Studies, Institute of Cybernetics at Tallinn University of Technology.


Title: On mathematical modelling of waves in nerve fibres.

Abstract: Beside celebrated action potential models for explaining the nerve pulse propagation, the recent attention is turned also to modelling of mechanical waves in fibre walls i.e., in cylindrical biomembranes. The Heimburg-Jackson model (2005) with a special type of nonlinearities has opened a wide avenue of studies of deformation waves in biomembranes. Here we present the analysis of the improved Heimburg-Jackson model which corresponds to the original (well-posed) Boussinesq equation including the higher order mixed derivatives. In this case the double dispersion model describes correctly the velocity limits and accounts for inertia of lipids as microstructural constituents of biomembranes. Such a governing equation of longitudinal waves along the biomembrane admits the solitary wave solutions with the width controlled by the additional dispersion term. Using the ideas from the theory of rods, the transverse displacement is determined as the derivative (c.f. experiments by Tasaki, 1988). The analysis includes the description of all the solution types to such a governing equation, the demonstration of the emergence of trains of solitary waves from a general initial condition and the study of their interactions. An attempt is made to unite the physical phenomena of the action potential, the deformation wave in the surrounding biomembrane and the pressure wave in the axoplasm into one robust mathematical model. The coupling mechanism is proposed to be realized by additional forces.

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